The PhD program is offered jointly by the following institutions:

University of Coimbra

Faculty of Science and Technology (FCTUC)
Centre for Neuroscience and Cell Biology (CNC)

Technical University of Lisbon

Instituto Superior Técnico (IST)

New University of Lisbon

Faculty of Science and Technology (FCTUNL)
Institute of Chemical and Biologic Technology (ITQB)

University of Minho

School of Engineering (UMinho)